Narakusumo, Raden Pramesa, and Michael Balke. “FOUR NEW SPECIES OF EPHOLCIS WATERHOUSE, 1875 (COLEOPTERA: SCARABAEIDAE: MELOLONTHINAE: MAECHIDIINI) FROM THE MOLUCCAS, INDONESIA”. Treubia 46 (March 7, 2025): 35–50. Accessed December 6, 2025. https://ejournal.brin.go.id/treubia/article/view/9394.